ANNEX I HABITAT DESCRIPTIONS

 

In Ireland’s marine and coastal zone SACs have been designated under the Habitats Directive for seven marine and 16 coastal habitat types. Of these, one marine and four coastal habitats are defined by the Habitats Directive as priority habitats. Priority habitats are deemed to be habitats in danger of disappearance, in view of the proportion of their natural range within the EU. Member states have a particular responsibility for their conservation.

 

There are a further 12 Annex I habitats that occur in coastal SACs but they are not purely coastal. Click here for a list of these.

 

Detailed descriptions of all Annex I habitats can be found on the website of the UK Joint Nature Conservation Committee (JNCC), here.

 

A general description and classification of Irish habitat types, and their relationship to Annex I habitats, can be found in the Heritage Council publication A Guide to Habitats in Ireland. For habitats in the marine and coastal this can be accessed by clicking the following links:

 

Coastland Habitats

Peatlands

Exposed Rock/Disturbed Ground

Marine Subtidal Habitats

Heath and Dense Bracken

 

Marine Intertidal Habitats

Grassland and Marsh

 

Marine Water Body Habitats

Woodland and Scrub

 

 

 

 

Marine Habitats

 

Code

Habitat Name

Code

Habitat Name

1110

Sandbanks which are slightly covered by sea water all the time

1160

Large shallow inlets and bays

1130

Estuaries

1170

Reefs

1140

Mudflats and sandflats not covered by seawater at low tide

8330

Submerged or partially submerged sea caves

1150

Coastal lagoons

 

 

 

Sandbanks which are slightly covered by sea water all the time (Habitat Code 1110)

These consist of sandy sediments that are permanently covered by shallow sea water, typically at depths of less than 20m. Sandbanks are distinct elongated, rounded or irregular ‘mound’ shapes of sandy sediment. The animal and plant communities associated with this habitat are determined primarily by sediment type, but also by factors such as wave exposure, depth, turbidity and salinity of the surrounding water. They are typically colonised by burrowing fauna, e.g. bivalve molluscs, worms and crustaceans. On the surface of the sandbank shrimps, gastropod molluscs, crabs and fish occur. Sandbanks are often important nursery areas for fish, and, due to the presence of sand-eels, they act as feeding grounds for seabirds and sea-duck. Sandbanks are frequently associated with other marine Annex I habitats; for example they may be a component habitats of Estuaries and Large shallow inlets and bays.

 

To date just one SAC has been designated due to the presence of Sandbanks – Long Bank (Co. Wexford). Sandbanks is one of the qualifying habitats in the Lower Shannon SAC.

 

Estuaries (Habitat Code 1130)

This habitat encompasses the downstream part of a river valley where the river enters the sea and is subject to the tide. Freshwater and seawater mix together and the decrease in current results in the deposition of fine sediments – often forming extensive intertidal sand and mudflats. Benthic algae and invertebrates flourish and provide a rich food source for wading birds. Many estuaries are also designated as SPAs under the Birds Directive.

 

Dundalk Bay (Louth), Rogerstown Estuary (Dublin), Bannow Bay (Wexford), Castlemaine Harbour (Kerry) and Lower River Shannon (Clare/Limerick/ Kerry) are important examples of estuaries designated as SACs.

 

Mudflats and sandflats not covered by seawater at low tide (Habitat Code 1140)

This habitat type can be divided into three broad categories - clean sands, muddy sands and muds. In practice there is a continuous gradation between them. The plant and animal communities present vary according to the type of sediment, its stability and the salinity of the water. They usually have no vascular plants but may have rich communities of blue algae, diatoms, eelgrass and invertebrates. As with estuaries, they are of particular importance as feeding areas for wildfowl and waders and are often also designated as SPAs.

 

Examples of intertidal mud/sandflat SACs include Malahide Estuary, Baldoyle Bay and North Dublin Bay (Dublin); Broadhaven Bay and Killala Bay/Moy Estuary (Mayo); and Tralee Bay (Kerry).

 

Coastal lagoons (Habitat Code 1150)

This priority habitat category encompasses areas of shallow, coastal salt water that are wholly or partially separated from the sea by sandbanks, shingle or, less frequently, rocks. The water in lagoons can vary in salinity from hypersaline (i.e. more salty than seawater as a result of evaporation) to brackish (owing to dilution of seawater by freshwater). Water levels in coastal lagoons may also vary considerably. The plant and animal communities of lagoons vary according to the physical characteristics and salinity regime of a particular lagoon. The vegetation may include beds of eelgrass, tasselweed, and pondweeds, or stoneworts. In more rocky lagoons, communities of brown (wracks and kelps), red and green algae are also found. The fauna is often characterised by mysid shrimps and other small crustaceans, worms that burrow into the sediment, gastropod molluscs, and some fish species. For further information on coastal lagoons in Ireland and a full list of sites see Healy (2003).

 

Cross Lough (Mayo), Dunbeacon Shingle (Cork), Lady’s Island Lake (Wexford) and Tacumshin Lake (Wexford) are SACs with excellent examples of coastal lagoons.

 

Large shallow inlets and bays (Habitat Code 1160)

Large shallow inlets and bays are large indentations of the coast, predominantly under the influence of salt water but generally more sheltered from wave action than the open coast. They are relatively shallow; with water less than 30m over most of the area.

 

They are effectively habitat complexes that comprise an interdependent mosaic of subtidal and intertidal habitats, many of which are also Annex I habitats (e.g. reefs, sandbanks and mudflats). Consequently, they contain a great diversity of sediments (from mud to sands to rock) and substrates with highly diverse animal and plant communities. The issue of site size is also important, as larger sites tend to encompass the greatest variety of constituent habitats and have the greatest potential for maintenance of ecosystem integrity.

 

Some of the largest and best-known bays along the south-west, west and north-west coasts are designated SACs. These include Roaringwater Bay (Cork), Kenmare Bay (Cork/Kerry), Kilkieran Bay (Galway), Clew Bay (Mayo) and Mulroy Bay (Donegal).

 

Reefs (Habitat Code 1170)

Reefs rise from the seabed and are either rocky marine habitats (i.e. rock, boulders and cobbles) which support animal and plant communities or biogenic reefs (where the structure is created by the animals themselves, e.g. cold-water coral reefs). Reefs are generally subtidal but may extend as an unbroken transition into the intertidal zone. Intertidal areas are only included within this Annex I type where they are connected to subtidal reefs.

 

Rocky reefs are extremely variable, both in form (rock walls to horizontal ledges, bed rock, boulder fields, and cobbles) and in the communities they support. Their communities are characterised by attached algae (where there is sufficient light) and invertebrates, usually associated with a range of mobile invertebrates and fish. The specific communities that occur are determined by a number of factors, for example, rock type and topography, wave and tidal exposure, turbidity and salinity. On intertidal reefs a strong vertical zonation is apparent, lichens occur at the top of the shore, grading into communities characterised by barnacles, mussels or species of fucoid (wrack) seaweeds.

The main species which form biogenic reefs in Irish waters are blue mussels (Mytilus edulis), horse mussels (Modiolus modiolus), ross worms (Sabellaria spp.), and cold-water corals such as (Lophelia pertusa).

 

Important examples of SACs designated for reef habitats include Wicklow Reef, Kilkee Reef (Clare), Kerry Head Shoal (Kerry), Carnsore Point (Wexford) and the Magharee Islands (Kerry).

 

To date, none of the Special Areas of Conservation designated by Ireland under the Habitats Directive extend beyond Ireland's territorial waters (12 nautical mile limit). The National Parks and Wildlife Service of the Department of Environment, Heritage and Local Government is currently in the process of selecting cold-water coral reef sites that are of conservation importance for designation as SACs.

 

Submerged or partially submerged sea caves (Habitat Code 8330)

This Annex I habitat category includes submerged sea caves and partially submerged caves which are only exposed to the sea at high tide. They are typically associated with the Annex I habitat Reefs. The animal and plant communities in sea caves vary considerably depending on the structure and extent of the cave, their degree of submergence and exposure to sand scour and wave-surge. Intertidal and shallow subtidal caves are often subject to conditions of strong wave surge and have floors of coarse sediment, cobbles and boulders. They are generally characterised by communities of mussels, barnacles, and encrusting animal species (e.g. sponges). In deeper water, caves are subject to less water movement and silt may accumulate on the cave floor. Such caves may provide shelter for crabs, lobsters and fish such as gobies.

 

Good examples of this habitat can be found in the following SACs – Lough Hyne Nature Reserve (Cork), Saltee Islands (Wexford); Blasket Islands (Kerry); Black Head Poulsallagh Complex (Clare) and Inishmore Island (Galway).

 

 

 

Coastal Habitats

 

Code

Habitat Name

Code

Habitat Name

1210

Annual vegetation of drift lines

2110

Embryonic shifting dunes

1220

Perennial vegetation of stony banks

2120

Shifting dunes along the shoreline with Ammophila arenaria (white dunes)

1230

Vegetated sea cliffs of the Atlantic and Baltic coasts

2130

Fixed coastal dunes with herbaceous vegetation (grey dunes)

1310

Salicornia and other annuals colonizing mud and sand

2140

Decalcified fixed dunes with Empetrum nigrum

1320

Spartina swards

2150

Atlantic decalcified fixed dunes

1330

Atlantic salt meadows

2170

Dunes with Salix repens ssp argentea

1410

Mediterranean salt meadows

2190

Humid dune slacks

1420

Mediterranean and thermo-Atlantic halophilus scrubs

21A0

Machairs

 

 

Annual vegetation of drift lines (Habitat Code 1210)

This habitat type occurs on deposits of small shingle lying at or above mean high-water spring tides. These shingle deposits occur as fringing beaches that are subject to periodic displacement or overtopping by high tides and storms. The sparse cover of distinctive vegetation is composed of annual or short-lived perennial species.

 

Plant species are also tolerant of saltwater inundation, as the beaches are often over-topped by the tide or subject to spray from waves breaking over the beach, e.g. Cakile maritima.

 

The SACs at Bannow Bay (Wexford), Castlemaine Harbour (Kerry), Clonakilty Bay (Cork), Tramore Dunes and Backstrand (Waterford) and Tranarossan and Melmore Lough (Donegal) all contain good examples of this habitat.

 

Perennial vegetation of stony banks (Habitat Code 1220)

This habitat consists of coastal areas where shingle (cobbles and pebbles) has accumulated to form elevated ridges or banks above the high tide mark. These areas provide a substrate for perennial vegetation, including Sea Sandwort, Sea Beet, Rock Samphire and Sea Mayweed. The rare perennial plant Sea-kale (Crambe maritima) can also be found on shingle. The strandline is excluded from this habitat as it is considered to be a separate habitat (Annual vegetation of drift lines).

 

Examples of SACs with this habitat include Cross Lough (Mayo), Tramore Dunes and Backstrand (Waterford), Lady’s Island Lake (Wexford), Dunbeacon Shingle (Cork) and Reen Point Shingle (Cork).

 

Vegetated sea cliffs of the Atlantic and Baltic coasts (Habitat Code 1230)

This category of habitats encompasses a complex range of cliff micro-habitats depending upon geology, geomorphology and the degree of exposure. It includes cliff crevices, ledges and slopes and extends to cliff tops with heaths, grasslands and scrub.

 

Examples of SACs with this habitat include Slieve League (Donegal), Mount Brandon (Kerry), Hook Head (Wexford), Howth Head (Dublin) and Saltee Islands (Wexford).

 

Salicornia and other annuals colonizing mud and sand (Habitat Code 1310)

This pioneer saltmarsh vegetation colonises intertidal mud and sandflats in areas protected from strong wave action. It is an important precursor to the development of more stable saltmarsh vegetation (e.g. habitat codes 1320, 1330, 1410 and 1420). Salt marshes usually comprise the upper, vegetated portions of intertidal mudflats, lying approximately between mean high water neap tides and mean high water spring tides. They are usually associated with estuaries and coastal lagoons. This particular salt marsh habitat is composed predominantly of glassworts Salicornia spp. with some grass and seablite species. For further information on saltmarshes in Ireland and a full list of sites see Curtis and Sheehy-Skeffington (1998) and Curtis (2003).

 

Examples of SACs with this habitat include Lackan Saltmarsh and Kilcummin Head (Mayo), River Barrow and River Nore (Waterford/Wexford), North Dublin Bay and Courtmacsherry Estuary (Cork).

 

Spartina swards (Spartinion maritimae) (Habitat Code 1320)

This salt marsh habitat refers to the perennial pioneer grasslands of coastal salt muds, formed by cord-grasses Spartina or similar grasses. Spartina is an alien species to Ireland, which has colonised rapidly. It is considered by some to be a threat to nature conservation and in some areas attempts have been made to control its spread. It can rapidly colonise mud and sandflats to the perceived detriment of other communities. However, it also has beneficial impacts on its habitat. For further information on Spartina in Ireland see McCorry et al. (2003). For further information on saltmarshes in Ireland and a full list of sites see Curtis and Sheehy-Skeffington (1998) and Curtis (2003).

 

Examples of SACs with this habitat include Lough Swilly (Donegal), Bannow Bay (Wexford), Rogerstown Estuary (Dublin) and Great Island Channel (Cork) (Cork).

 

Atlantic salt meadows (Glauco-Puccinellietalia maritimae) (Habitat Code 1330)

This salt marsh habitat forms in sheltered areas within estuaries, e.g. behind sand spits and in shallow protected bays, where tidal inundation still occurs but with decreasing frequency and duration. A wide range of community types are found and the saltmarshes can cover large areas, especially where there has been little or no enclosure on the landward side. Important plant species include Common saltmarsh-grass (Puccinellia maritima), Sea purslane (Halimione pedunculata) and sea aster (Aster tripolium). For further information on saltmarshes in Ireland and a full list of sites see Curtis and Sheehy-Skeffington (1998) and Curtis (2003).

 

Examples of SACs with this habitat include Lackan Saltmarsh and Kilcummin Head (Mayo), Bellacragher Saltmarsh (Mayo), Streedagh Point Dunes (Sligo), Raven Point Nature Reserve (Wexford) and Courtmacsherry Estuary (Cork).

 

Mediterranean salt meadows (Juncetalia maritimi) (Habitat Code 1410)

This salt marsh habitat occurs in the uppermost levels of saltmarshes. It is subject to less frequent and less prolonged innundation by the sea. Vegetation is typically dominated by rushes and sedges, e.g. Juncus maritimus, J. gerardii and Carex extensa. For further information on saltmarshes in Ireland and a full list of sites see Curtis and Sheehy-Skeffington (1998) and Curtis (2003).

 

Examples of SACs with this habitat include Bellacragher Saltmarsh (Mayo), The Murrough Wetlands (Wicklow), Lacken Saltmarsh and Kilcummin Head (Mayo), and Sheephaven (Donegal).

 

Mediterranean and thermo-Atlantic halophilus scrubs (Sarcocornetea fruticosi) (Habitat Code 1420)

This salt-tolerant scrubby vegetation, which is not common in Ireland, develops in the uppermost levels of saltmarshes, often where there is a transition from saltmarsh to dunes, or in some cases where dunes overlie shingle. Just two SACs, Ballyteigue Burrow and Bannow Bay (Co. Wexford), are designated due, in part, to the presence of this habitat. For further information on saltmarshes in Ireland and a full list of sites see Curtis and Sheehy-Skeffington (1998) and Curtis (2003).

 

Embryonic shifting dunes (Habitat Code 2110)

This habitat represents the early stage in sand dune development. Small mounds of sand (generally < 1 m in height) build up on the upper seashore, between the high tide mark and marram dunes, where wind-blown sand is impeded around plants on the drift-line (e.g. Sand Couch and Lyme-grass). The sand mounds are largely unvegetated but may support salt-tolerant plants such as Sea Rocket, Saltwort and Sea Sandwort.

 

This habitat type rarely occurs in isolation, because it may initiate dune succession, and it is invariably one of several Annex I habitat types to be found on a dune system. In many cases Embryonic shifting dunes will either be displaced by "white dunes" as the dunes develop or will be washed away by storms. Creation of new dune habitat is often dependent upon the survival of this habitat type.

 

This habitat is present in many of the major dune systems that area designated as SACs, e.g. Cahore Polders and Dunes (Wexford), Tramore Dunes and Backstrand (Waterford), Kilkeran lake and Castlefreke Dunes (Cork), Lough Nagreaney Dunes (Donegal) and Kilpatrick Sandhills (Wexford).

 

Shifting dunes along the shoreline with Ammophila arenaria (white dunes) (Habitat Code 2120)

This habitat represents the second stage in sand dune development – after embryonic dunes. They are partially stabilised hills or ridges of sand that occur along the seaward edge of the main sand dune system. Vegetation cover is incomplete and loose sand at the surface is subject to wind-blow. Tussocks of the familiar dune-building species Marram (Ammophila arenaria) are commonly interspersed with species such as Sand Sedge, Sea Spurge and Sea-Holly.

 

Good examples of white dunes are present in SACs at North Dublin Bay, Dog’s Bay (Galway), Gweedore Bay and Islands (Donegal), Magherabeg Dunes (Wicklow), Streedagh Point Dunes (Sligo) and Kilmuckridge-Tinnaberna Sandhills (wexford).

 

Fixed coastal dunes with herbaceous vegetation (grey dunes) (Habitat Code 2130)

This priority habitat is the final stage in the formation of sand dunes. The Ammophila dunes are succeeded by the fixed dune stage. Fixed dunes are stabilised ridges or hills of sand with a more or less complete cover of vegetation. Plant species composition is highly variable and the vegetation is usually characterised by herb-rich grassland or heath communities with extensive cover of mosses and lichens. 'Grey dunes' are so-called because of the abundance of grey-coloured lichens in the vegetation. Fixed dunes also include consolidated and flattened dune areas that can occur behind the main dune ridges. These areas are frequently used for agriculture, and grassland communities may contain 'agricultural' species such as Perennial Rye-grass, White Clover, Daisy and Dandelion.

 

Grey dunes invariably occur in many of the SACs that contain embryonic dunes and white dunes.

 

Decalcified fixed dunes with Empetrum nigrum (Habitat Code 2140)

This priority habitat represents the later, more mature, stages of sand dune succession. Exposure to rainfall over long periods results in leaching of the surface layers, causing a loss of calcium carbonate and increased soil acidity. The dunes become colonised by heath species such as crowberry (Empetrum nigrum) and ling heather (Calluna vulgaris). Decalcified fixed dunes often occur in mosaics with other habitats, depending on local physical and soil conditions, e.g. Atlantic decalcified fixed dunes, “grey dunes" and Humid dune slacks. Grazing helps to maintain the open nature of the vegetation. Undergrazing would result in the development of scrub and woodland through the process of succession. However, fixed dunes are also vulnerable to overgrazing.

 

Just five SACs are designated due to the presence of this priority habitat, all in County Donegal; Lough Nagreaney Dunes, Slieve Tooey/Tormore Island/Loughros Beg Bay, Tranarossan and Melmore Lough, West of Ardara/Maas Road and Gweedore Bay and Islands.

 

Atlantic decalcified fixed dunes (Calluno-Ulicetea) (Habitat Code 2150)

This priority habitat, which occurs on mature, stable dunes where the initial calcium carbonate content of the dune sand is low, is similar in composition to Decalcified fixed dunes with Empetrum nigrum. At most sites at which it occurs, it forms a mosaic with other Annex I habitat types. It can be difficult to distinguish between these two Annex I habitat types, as the vegetation forms a continuous spectrum of variation within complex habitat mosaics. The most characteristic community is dune heath, with ling heather Calluna vulgaris found in combination with sand sedge Carex arenaria.

 

Atlantic decalcified fixed dunes is more widespread than Decalcified fixed dunes with Empetrum nigrum (which is restricted to the north-west coast) as it tolerates drier and warmer conditions. This is reflected in the distribution of designated SACs that include this habitat; e.g. Mweelrea/Sheeffry/Erriff Complex (Mayo), Buckroney-Brittas Dunes and Fen (Wicklow), Ballyteigue Burrow (Wexford), Clonakilty Bay (Cork) and Lough Nagreaney Dunes (Donegal).

 

Dunes with Salix repens ssp argentea (Salix arenariae) (Habitat Code 2170)

This habitat comprises dunes or parts of dunes where creeping willow (Salix repens ssp. Argentea) is dominant, forming prominent, low scrubby growth. Creeping willow grows predominantly in and around dune slacks, though on some sites it may spread up the drier ridges. This habitat often occurs alongside Humid dune slacks and the boundaries between the two are often diffuse and difficult to define on the ground.

 

Good examples of this habitat are present in SACs at Horn Head and Ringclevan (Donegal), Castlemaine Harbour (Kerry), Gweedore Bay and Islands (Donegal), Mweelrea/Sheeffry/Erriff Complex (Mayo) and Raven Point Nature Reserve (Wexford).

 

Humid dune slacks (Habitat Code 2190)

Dune slacks are low-lying areas within dune systems that are occasionally flooded and where nutrient levels are low. The range of communities found is considerable and depends on the structure of the dune system, the successional stage of the dune slack and variations in the extent and duration of flooding of the dune surface. Creeping willow is often found in dune slack vegetation, although it is not dominant; as it is in Dunes with Salix repens ssp. Argentea. Dune slacks are often rich in plant species, particularly rare and local species, e.g. the Annex II species Petalwort (Petalophyllum ralfsii).

 

Good examples of this habitat are present in SACs at Ballyness Bay (Donegal), Akeragh, Banna and Barrow Harbour (Kerry), Ballysadare Bay (Sligo), Buckroney-Brittas Dunes and Fen (Wicklow) and North Dublin Bay.

 

Machairs (Habitat Code 21A0)

This priority habitat consists of coastal grassy plains that are formed of wind-blown calcareous sands. The plains are usually almost level, however, they may also have an undulating or hillocky surface, and can even occur on hills. They develop in places with a cool, moist and windy climate. In Ireland, they are limited to the west coast from Galway Bay to Malin Head, in Donegal. Machair vegetation is often species-rich and features elements of sand dune communities and calcareous grassland.

 

Machair areas were often widely cultivated in the past but now many support extensive grazing regimes and may be used for amenity (sports pitches, camp sites, caravan parks) or development. The sandy substrate tends to be unstable and may be disturbed by winter gales, especially if grazing is heavy. Machairs often support large breeding bird populations and is particularly important for waders and corncrake.

 

Other Habitats

 

Code

Habitat Name

Code

Habitat Name

4010

Northern Atlantic wet heaths with Erica tetralix

6510

Lowland hay meadows

4030

European dry heaths

7130

Blanket bog

4060

Alpine and Boreal heaths

8210

Calcareous rocky slopes with chasmophytic vegetation

5130

Juniperus communis formations on heaths or calcareous grasslands

8220

Siliceous rocky slopes with chasmophytic vegetation

6210

Semi-natural dry grasslands and scrubland facies on calcareous substrates

8240

Limestone pavements*

6410

Molinia meadows on calcareous, peaty or clavey-silt-laden soils

91A0

Old sessile oak woods with Ilex and Blechnum in British Isles

 

Descriptions of all of these habitats can be seen on the JNCC Annex I habitat accounts site, here.

 

 

 


URLs in Text

JNCC Habitats Pages - http://www.jncc.gov.uk/ProtectedSites/SACselection/SAC_habitats.asp

HC Habitats Book - http://heritagecouncil.ie/publications/habitats/

Coastland Habitats - http://www.heritagecouncil.ie/publications/habitats/11.html

Marine Subtidal Habitats- http://www.heritagecouncil.ie/publications/habitats/13.html

Marine Intertidal Habitats - http://www.heritagecouncil.ie/publications/habitats/12.html

Marine Water Body Habitats - http://www.heritagecouncil.ie/publications/habitats/14.html

Peatlands - http://www.heritagecouncil.ie/publications/habitats/7.html

Heath and Dense Bracken - http://www.heritagecouncil.ie/publications/habitats/7.html

Grassland and Marsh - http://www.heritagecouncil.ie/publications/habitats/5.html

Woodland and Scrub - http://www.heritagecouncil.ie/publications/habitats/8.html

Exposed Rock/Disturbed Ground - http://www.heritagecouncil.ie/publications/habitats/9.html

Curtis, T.G.F. and Sheehy Skeffington, M. (1998) - http://www.ria.ie/publications/journals/procbi/1998/PB98I2/PDF/98208BI.pdf

 

 

Other URLs of Interest

UK Habitat Action Plans - http://www.ukbap.org.uk/habitats.aspx

Link between general habitats and Anex I habitats in Ireland - http://www.heritagecouncil.ie/publications/habitats/app1.html

Heritage Council Study of Impacts on Designated Conservation Areas - http://heritagecouncil.ie/publications/coastalrep/index.htm

 

 

Selected Further Reading

Bassett, J. A. 1983 Report on the Conservation of Irish Coastal Sites. Machair in Ireland. Unpublished Report. Wildlife Service. Dublin.

Bassett, J.A. and Curtis, T.G.F. (1985). The nature and occurrence of sand-dune machair in Ireland. Proceedings of the Royal Irish Academy, 85B (1): 1-20.

Curtis, T.G.F. (1991). A site inventory of the sandy coasts of Ireland. In: Quigley, M.B. (ed) A Guide to the Sand Dunes of Ireland. 3rd Congress of the European Union for Dune Conservation and Coastal Management, Galway.

Curtis, T.G.F. (2003). Saltmarshes. In: Otte, M.L. (ed.) Wetlands of Ireland - Distribution, Ecology, Uses and Economic Value. UCD Press. pp. 31-43

Curtis, T.G.F. and Sheehy Skeffington, M. (1998). The Salt Marshes of Ireland: An inventory and account of their geographical variation. Biology and Environment: Proc. R. Ir. Acad., 98B (2): 87-104. http://www.ria.ie/publications/journals/procbi/1998/PB98I2/PDF/98208BI.pdf

European Commission (1999). Interpretation Manual of European Habitats. EUR 15/2. DG Environment, European Commission.

Healy, B. 1994 Lagoons and other enclosed brackish waters in the Republic of Ireland. Dept. Zoology. UCD

Healy, B. (2003). Coastal Lagoons. In: Otte, M.L. (ed.) Wetlands of Ireland - Distribution, Ecology, Uses and Economic Value. UCD Press. pp. 51-78.

McCorry, M, Curtis, T.G.F. and Otte, M.L. (2003). Spartina in Ireland. In: Otte, M.L. (ed.) Wetlands of Ireland - Distribution, Ecology, Uses and Economic Value. UCD Press. pp. 44-50.

Moore, D. and Wilson, F. (1999). The National Shingle Beach Survey of Ireland. Unpublished Report, NPWS, Dublin.

Neff, J. (1998). Irish Coastal Habitats: A Study of Impacts on Designated Conservation Areas. The Heritage Council, Kilkenny. http://heritagecouncil.ie/publications/coastalrep/index.htm

Otte, M.L. (2003). Wetlands of Ireland - Distribution, Ecology, Uses and Economic Value. UCD Press.

Quigley, M.B. (1991) A Guide to the Sand Dunes of Ireland. 3rd Congress of the European Union for Dune Conservation and Coastal Management, Galway.